Table 3 OSR and parameters of male and female distribution at female vernal emergence for subsequent breeders and nonbreeders.

Data are presented as mean ± SD [range] (n) (see statistics in Table 1).

ParametersBreedersNonbreeders
YearlingsFemales aged ≥2 yearsYearlingsFemales aged ≥2 years
Mean distance to the five
nearest male burrows, m		95 ± 54 [26–236] (18)106 ± 57 [42–251] (38)152 ± 60 [29–230] (12)105 ± 34 [59–251] (16)
Number of male* burrows within 100 m
of the female’s burrow		3.2 ± 2.3 [0–6] (18)3.1 ± 1.7 [0–6] (38)1.7 ± 1.6 [0–6] (12)2.4 ± 1.4 [1–5] (16)
Mean distance to the five nearest
female burrows, m		50 ± 29 [26–128] (18)66 ± 61 [18–352] (38)50 ± 26 [23–95] (12)59 ± 30 [18–109] (16)
Number of female† burrows within 70 m
of the female’s burrow		3.4 ± 3.1 [0–9] (18)3.7 ± 3.1 [0–12] (38)2.5 ± 2.2 [0–8] (12)2.9 ± 2.3 [0–9] (16)
OSR (female/male)0.6 ± 0.3 [0.2–1.1] (27)0.7 ± 0.4 [0.08–1.3] (47)0.9 ± 0.3 [0.6–1.2] (12)1.0 ± 0.3 [0.2–1.3] (19)

*Only ≥2-year-old males that emerged no later than the fifth day after the date of emergence of the focal female.

†Only females that emerged within ±5 days from the date of emergence of the focal female.